Brushes become a lighter tint of black in an RGB document when expanded or rasterized.
Brushes like Art Brushes are based on a CMYK document. They were created with C0 M0 Y0 K100.
When these brushes are used in an RGB document with an RGB black (0/0/0) for the stroke, the result will be around a lighter 10% black than expected because the CMYK black will be converted to RGB.
You can check that by expanding, rasterizing or Output Preview in Acrobat.
If they had been created in the CMYK document with a 100% Grey instead of CMYK values, they would have converted to the darkest black in an RGB documents (greyscale always keep their values when converted between RGB and CMYK documents).
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Ton commented
I could not believe I could not find it reported. I comes up from time to time during the years at the Illustrator forum.
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I was sure a report for this exists... but I bet it’s a false memory, based on many Von’s videos mentioning it
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Ton commented
Here is an example set.